The TDG protein environment connects active DNA demethylation with chromatin and RNA biology

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Argitaratua izan da:Cellular and Molecular Life Sciences vol. 83, no. 1 (Dec 2026), p. 15
Egile nagusia: Richina, Federica
Beste egile batzuk: Noreen, Faiza, Bauer, Christina, Weber, Alain, Kunz, Christophe, Buczak, Katarzyna, Schwarz, Simon D., Wu, Fabian, Schürmann, David, Schär, Primo
Argitaratua:
Springer Nature B.V.
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Sarrera elektronikoa:Citation/Abstract
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100 1 |a Richina, Federica  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
245 1 |a The TDG protein environment connects active DNA demethylation with chromatin and RNA biology 
260 |b Springer Nature B.V.  |c Dec 2026 
513 |a Journal Article 
520 3 |a Understanding the interplay between DNA methylation and gene expression remains a challenge. This study explores the proteome of active DNA demethylation in murine embryonic stem cells (mESC), focusing on the base-excision-repair (BER) step initiated by the Thymine DNA Glycosylase (TDG). Using BioID2 proximity labeling, we identified a TDG interactome encompassing four functional aspects: chromatin organization and transcription, chromosomal organization, RNA processing, and ribosomal biogenesis. We show specifically that TDG participates in a genome regulatory network involving chromatin remodelers and modifiers such as RUVBL2 and the H3K4 methyltransferase complex tethering factor HCFC1, consistent with the dysregulation of histone modifications observed in TDG-deficient cells. We also identified the paraspeckle components PSPC1 and NONO as TDG interactors, implicating TDG in RNA-mediated nuclear processes. This led us to show that TDG is an RNA-binding protein, interacting with long-noncoding RNAs (lncRNA), including the paraspeckle organizing lncRNA Neat1, previously reported to target TET proteins to genomic sites and to engage in R-loop regulation. We then demonstrate TDG’s ability to excise oxidized 5-methylcytosine in RNA:DNA hybrids, suggesting a role of active DNA demethylation in the regulation of R-loops. Our findings thus unveil a direct crosstalk between active DNA demethylation, chromatin modification and remodeling as well as RNA-genome interactions in mESC, providing avenues for future mechanistic investigations. 
653 |a DNA methylation 
653 |a rRNA 
653 |a Embryo cells 
653 |a Tethering 
653 |a Methyltransferase 
653 |a Thymine 
653 |a Humidity 
653 |a Histones 
653 |a Ribonucleic acid--RNA 
653 |a Stem cells 
653 |a DNA glycosylase 
653 |a Chromatin remodeling 
653 |a Genomes 
653 |a Non-coding RNA 
653 |a Epigenetics 
653 |a Hybrids 
653 |a Demethylation 
653 |a Proteins 
653 |a Plasmids 
653 |a Oxidation 
653 |a Thymine DNA glycosylase 
653 |a Functional morphology 
653 |a Gene expression 
653 |a RNA-binding protein 
653 |a Cloning 
653 |a Deoxyribonucleic acid--DNA 
653 |a Proteomes 
653 |a RNA processing 
653 |a R-loops 
653 |a Environmental 
700 1 |a Noreen, Faiza  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642); Swiss Institute of Bioinformatics, Basel, Switzerland (GRID:grid.419765.8) (ISNI:0000 0001 2223 3006) 
700 1 |a Bauer, Christina  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Weber, Alain  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Kunz, Christophe  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Buczak, Katarzyna  |u University of Basel, Proteomics Facility Biozentrum, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Schwarz, Simon D.  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Wu, Fabian  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Schürmann, David  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
700 1 |a Schär, Primo  |u University of Basel, Department of Biomedicine, Basel, Switzerland (GRID:grid.6612.3) (ISNI:0000 0004 1937 0642) 
773 0 |t Cellular and Molecular Life Sciences  |g vol. 83, no. 1 (Dec 2026), p. 15 
786 0 |d ProQuest  |t Health & Medical Collection 
856 4 1 |3 Citation/Abstract  |u https://www.proquest.com/docview/3288173756/abstract/embedded/7BTGNMKEMPT1V9Z2?source=fedsrch 
856 4 0 |3 Full Text  |u https://www.proquest.com/docview/3288173756/fulltext/embedded/7BTGNMKEMPT1V9Z2?source=fedsrch 
856 4 0 |3 Full Text - PDF  |u https://www.proquest.com/docview/3288173756/fulltextPDF/embedded/7BTGNMKEMPT1V9Z2?source=fedsrch